During weight maintenance a high saturated fat, high beef diet increased the number of small dense LDL particles.

In a study co-authored by Ronald Krauss, during weight maintenance two low-carb diets (one high and one low in saturated fat) were compared to a high-carb diet that was also high in saturated fat. The values for small dense LDL were 222 nmol/l in the low-carb, high saturated fat diet ; 207 nmol/l in the high-carb diet, high saturated fat diet; and 187 nmol/l in the low-carb, low saturated fat diet.

Mangravite LM, Chiu S, Wojnoonski K, Rawlings RS, Bergeron N, Krauss RM. Changes in atherogenic dyslipidemia induced by carbohydrate restriction in men are dependent on dietary protein source. J Nutr.2011Dec;141(12):2180-5.
http://www.slideshare.net/pronutritionist/beef-low-carb-diet (see slide 6)

What Dr. Krauss said about this study:
"This is one of the surprises that keep life interesting for us for us as researchers and also for the world out there who happens to be looking over our shoulder. In this case, the surprise was that the combination of the high beef diet and the high saturated fat diet caused very serious increases in all of the cholesterol related risk factors we had been measuring, including total particle numbers, small LDL, total LDL cholesterol, inflammation, whatever we looked at, we saw an adverse effect."
http://www.meandmydiabetes.com/2012/04/17/ron-krauss-saturated-fat-red-meat-it-depends